Integrated ad blocker, enabled by default. It’s the one thing Firefox can do that Chrome never will.
Most people aren’t tech-savvy enough to install an ad blocker. Most people really do see all the ads on the Internet. Blocking ads by default would make the browsing experience far more pleasant, and so browsing with Firefox would be far more pleasant than browsing with Chrome for most people.
And obviously, continue to invest in improving the core web experience instead of getting distracted by side quests and firing everybody working on Servo, etc.