Spatial navigation is a feature I really do miss. I don't think any other browser supports this. It made keyboard-based browsing possible without resorting to stuff like hit-a-hint. You could just hit Shift+Arrow Key (which I mapped to the home row) and select a the nearest link (or anything interactive) in that direction. I think it worked in a visual fashion so order in the DOM didn't matter at all. It behaves exactly like one would expect.
The Opera CTO is now building Vivaldi, which is basically Opera. It has all the features I remember (spatial navigation, mail/RSS client, gestures, split tabs, etc) and is very good.