With the app I tried to provide additional access points into the data. Page links are an additional way if you use them like "@ mentions". Let's say you have a person or topic that a note is related to (for an email this would be the sender). I created a folder "Contacts" and added a note for each person. Now if I write something down I mention the page for that person on the note. Later on I can go to that person's page and see all notes connected to that person (the power of backlinks) or simply search for "@james", for example.
Zettelkasten: I'm still trying to wrap my head around this one. What is great about it is that Niklas Luhmann figured out a way to have page links and backlinks between notes in a setup in the physical world on actual paper. With digital notes, I think GUIDs and timestamps to identify notes are not necessary anymore, because the file path (or URL if you will) uniquely identifies a note.
Org mode: yes, same here with the added difficulty that I have a vi background :)