I thought the success of YouTube was largely due to their low bars for quality of content. Put simply, they allow all kinds of crap as long as it generates views.
Vimeo's model is quality content from indie producers, actually they are similar to SoundCloud in some ways.
Think about this: for example NASA has plenty of quality audio and video materials, where should they upload it? SoundCloud and Vimeo would be the best choice and it's what they do. It's quality content not exactly "publishable" via music labels or television, and yet it's pretty good.
Not everyone wants to be on the Internet's sewage system that's YouTube, and not everyone wants to go the costly publishing route. There has to be a niche for this type of media. I'm sure monetization can be figured out when there's clearly a market for it.