It inverts the direction (up swaps with down and left swaps with right) from the previous trackpad scrolling behavior.
Before 10.7, the scroll behavior acts like you've grabbed the scroll bar and your fingers drag it, so moving your fingers down makes the scroll grip move down and the page content move up. Moving your fingers left makes the scroll grip move left and the page content move right. The scrolling paradigm is that your fingers grab the viewport and move it with respect to the content.
In 10.7 and later, it instead acts as though your fingers are grabbing the content and moving it around within a stationary viewport. So moving your fingers up pushes the content upward. So the scroll interaction is like the direct interaction on a touchscreen, but via a trackpad instead.