Before it ran on F1/Spanner, Adwords ran on sharded MySQL on Borg. Not for its entire early life, but for quite a few years. Later in life, Checkout and maybe Wallet ran on MySQL on Borg, too. So did YouTube, which used Vitess, a sharding layer (now ported to Kubernetes and open sourced).
Bigtable, Spanner and even Colossus/D run in containers on Borg.